After joining the youth setup of his local club Marila Příbram at the age of 10, Slepička broke into the first team in 1999. Finding his playing time limited, however, Slepička moved to Slovan Liberec and then to giants Sparta Prague, for whom he wore the prized number 10 jersey. He has had successful spells at both Sparta Prague and Croatian giants Dinamo Zagreb, winning trophies at both clubs as well as appearing in the Champions League for both. He has also made two appearances for the Czech Republic national team.
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Czech Republic